Website CV


  • Quantitative methods in fish and wildlife biology including biotelemetry data analysis, survival analysis, population estimation techniques, computer applications, large mammal population dynamics. Workshops on Program MARK.


Ph.D., Zoology - Ohio State University
Columbus, Ohio, 1976

M.S., Wildlife Biology - University of Maine
Orono, Maine, 1972

B.S., Fisheries and Wildlife Biology - Iowa State University
Ames, Iowa, 1970