Gary C. White

Professor Emeritus

Department of Fish, Wildlife and Conservation Biology
Colorado State University
Fort Collins, CO 80523
Office: (970)491-6678
FAX: (970)491-5091
Email: Gary.White@ColoState.edu

Retirement.

On June 30, 2007, I was emancipated from Colorado State University. This can be interpreted as meaning I plan to stay active professionally, but not take additional graduate students, and not continue serving on Faculty Council and the CSU ACUC. Currently, I'm still serving on graduate committees at Colorado State, consulting on various mark-encounter projects, conducting workshops on Program MARK, and traveling in our satellite-dish equipped Bigfoot motor home.

Research Interests.

Quantitative ecology, estimation of population parameters, compensatory mortality in mule deer populations, development of Program MARK.

Courses Taught.

I formally taught 2 graduate courses closely related to my research interests, FW662 Population Dynamics and FW 663 Analysis of Vertebrate Populations (with Paul F. Doherty, Jr.). In addition, I taught FW696, a graduate student orientation course each fall.

Recent past Ph. D. and M. S. student projects were:

bullet determining density-dependence in population growth (Tanya Shenk),
bullet determining strategies for when to winter feed mule deer (Bruce Lubow),
bullet evaluating the effects of archery season on elk movements (Mary Conner),
bullet movements of Preble's meadow jumping mouse, an endangered species (Arti Dharman),
bullet further evaluation of the effects of archery season on elk movements (Mark Post Vieira),
bullet population dynamics of mountain plover (Stephen J. Dinsmore),
bullet evaluating the impact of grazing on greater prairie chicken populations on the Valentine National Wildlife Refuge, Nebraska (Bridgette Flanders),
bullet estimating the population size of sage grouse in Middle Park, Colorado (Daniel Walsh),
bullet evaluating the effects on lesser prairie chicken populations of inter-seeding Conservation Reserve Program lands with native forbs in western Kansas (Tamara Fields),
bullet development of a population model of North Park Greater Sage-grouse populations (Kristen Strohm),
bullet effects of controlled burns on wildlife populations (Sarah J. Converse),
bullet synthesizing mark-recapture and harvest models to estimate demographic parameters (Paul Conn),
bullet estimation of mountain sheep populations in Rocky Mountain National Park (Brett McClintock),
bullet mule deer response to enhanced nutrition (Chad Bishop),
bullet improving efficiency of quadrat sampling to estimate elk populations (Greg Davidson), and
bullet habitat effects on snow shoe hare population dynamics (Jacob Ivan).

Software.

For a list of software that I have developed and how to obtain the code, see obtaining software. Most of my recent effort has been on the development of Program MARK.

Recent and Current Service Activities.

bullet Chaired a panel to review the USGS Wildlife Program and produce a written report report, October, 2009.
bullet Representative to The Wildlife Society Council for the Central Mountains and Plains Section November, 2008.
bullet Past-President, Central Mountains and Plains Section of The Wildlife Society, 2008-2009.
bullet Program MARK workshop in Las Cruces, New Mexico, April 6-11, 2008.
bullet Program MARK workshop in Riggins, Idaho, February 25-29, 2008.
bullet President, Central Mountains and Plains Section of The Wildlife Society, 2006-2008.
bullet Program MARK workshop in Fort Collins, Colorado, June 3-8, 2007, with Paul Doherty, Ken Burnham, and David Anderson.
bullet President-elect, Central Mountains and Plains Section of The Wildlife Society, 2005-2006.
bullet Faculty advisor to the Colorado State University Student Chapter of The Wildlife Society, 2002-2007.
bullet Associate Editor, Journal of Wildlife Management, 2006-present.
bullet Program MARK workshop in Perth, Australia, August 7-11, 2006, with David R. Anderson.
bullet Associate Editor, Wildlife Society Bulletin, 2002-2006.
bullet Program MARK workshop in the United Arab Emirates, June 22-28, 2005, with David R. Anderson.
bullet Program MARK workshop in Fort Collins, Colorado, June 5-10, 2005, with David R. Anderson and Kenneth P. Burnham.
bullet Program MARK workshop in Fort Collins, Colorado, June 7-11, 2004, with David R. Anderson and Kenneth P. Burnham.
bullet Program MARK workshop in Fort Collins, Colorado, June 2-6, 2003, with David R. Anderson and Kenneth P. Burnham.
bullet Program MARK workshop in Fort Collins, Colorado, June 3-7, 2002, with David R. Anderson and Kenneth P. Burnham.
bullet Statistical analysis of demographic data from California spotted owls, Fort Collins, Colorado, July 9-13, 2001.
bullet Program MARK workshop in Fort Collins, Colorado, June 4-8, 2001, with David R. Anderson and Kenneth P. Burnham.
bullet Program MARK workshop in Arcata, California, January 8-12, 2001, with Alan Franklin and Steve Dinsmore.
bullet Program MARK workshop in Fort Collins, Colorado, June 5-9, 2000, with David R. Anderson and Kenneth P. Burnham.
bullet Program MARK workshop in Fort Collins, Colorado, June 7-11, 1999, with David R. Anderson and Kenneth P. Burnham.
bullet Statistical analysis of demographic data from northern spotted owls, Corvallis, Oregon, December 7-14, 1998.
bullet Program MARK workshop in Dunedin, New Zealand, April 16-17, 1998, with Richard Barker.
bullet Program MARK workshop in St. Andrews, Scotland, August 10-14, 1998, with David R. Anderson and Kenneth P. Burnham.
bullet Guest of the South African Statistical Society, 29 October to 14 November, 1997. I presented lectures on population viability analysis, statistical modeling and parameter estimation, Program MARK, and mule deer density dependence.
bullet Program MARK workshop to Striped Bass Working Group, Reston, Virginia, June 16-10, 1997.
bullet Mexican Spotted Owl Recovery Team, 1993-present. My responsibilities involved developing delisting criteria and developing a population monitoring plan to meet the delisting criteria.
bullet Workshop in Erice, Sicily, Italy, 28 November to 3 December, 1996 on Research Techniques in Ethology and Animal Ecology: Uses and Misuses. I presented a lecture on population viability analysis.
bullet Associate Editor, Journal of Wildlife Management, 1993-1995.

Selected Professional Achievements.

bullet Elected representative to The Wildlife Society Council for the Central Mountains and Plains Section November, 2008.
bullet Elected President-elect of the Central Mountains and Plains Section of The Wildlife Society, August, 2005.
bullet 51st recipient of The Wildlife Society's Aldo Leopold Award and Medal for Distinguished Service to Wildlife Conservation, March 27, 2000.
bullet University of Maine Distinguished Wildlife Alumnus 1999.
bullet U. S. Department of Interior, Washington, D. C., 1999, Conservation Service Award to the Mexican Spotted Owl Recovery Team. Highest honor given to civilians by the Department of Interior.
bullet Colorado Chapter, The Wildlife Society 1988 Douglas L. Gilbert Award for Outstanding Professional Achievements in Wildlife Science.
bullet The Wildlife Society Monograph of the Year 1984.

Recent Publications.

White, G. C. 1996. NOREMARK: population estimation from mark-resighting surveys. Wildlife Society Bulletin. 24:50-52.

White, G. C., A. F. Reeve, F. G. Lindzey, and K. P. Burnham. 1996. Estimation of mule deer winter mortality from age ratios. Journal of Wildlife Management 60:37-44.

Burnham, K. P., D. R. Anderson, and G. C. White. 1996. Meta-analysis of vital rates of the Northern Spotted Owl. Studies in Avian Biology 17:92-101.

Bennetts, R. E., G. C. White, F. G. Hawksworth, and S. E. Severs. 1996. The influence of dwarf mistletoe on bird communities in Colorado ponderosa pine forests. Ecological Applications 6:899-909.

White, G. C., and R. E. Bennetts. 1996. Analysis of frequency count data using the negative binomial distribution. Ecology 77:2549-2557.

Lubow, B., G. C. White, and D. R. Anderson. 1996. Evaluation of a linked-sex harvest strategy for cervid populations. Journal of Wildlife Management 60:787-796.

Miller, S. G., G. C. White, et al. 1997. Brown and black bear density estimation in Alaska using radiotelemetry and replicated mark-resight techniques. Wildlife Monographs 133:1-55.

White, G. C. and R. M. Bartmann. 1997. Density dependence in deer populations. Pages 120-135 in The Science of Overabundance: Deer Ecology and Population Management. W. J. McShea, H. B. Underwood, and J. H. Rappole, eds. Smithsonian Institution Press, Washington, D.C.

White, G. C., and R. M. Bartmann. 1997. Effect of density reduction on overwinter survival of free-ranging mule deer fawns. Journal of Wildlife Management 62:214-225.

White, G. C., and R. M. Bartmann. 1998. Mule deer management – what should be monitored? Pages 102-116 in Journal C. Vos, Jr., editor. Proceedings of the 1997 deer-elk workshop, Rio Rico, Arizona. Arizona Game and Fish Department, Phoenix, Arizona, USA.

White, G. C., and R. M. Bartmann. 1998. Effect of density reduction on overwinter survival of free-ranging mule deer fawns. Journal of Wildlife Management 62:214-225.

Anderson, D. R., K. P. Burnham, and G. C. White. 1998. Comparison of Akaike information criterion and consistent Akaike information criterion for model selection and statistical inference from capture-recapture studies. Journal of Applied Statistics 25(2):263-282.

Shenk, T. M., G. C. White, and K. P. Burnham. 1998. Sampling-variance effects on detecting density dependence from temporal trends in natural populations. Ecological Monographs 86:445-463.

Thompson, W. L., G. C. White, and C. Gowan. 1998. Monitoring Vertebrate Populations. Academic Press, New York, NY. 365 pp.

Unsworth, J. W., D. F. Pac, G. C. White, and R. M. Bartmann. 1999. Mule deer survival in Colorado, Idaho, And Montana. Journal of Wildlife Management 63:315-326.

White, G. C., and K. P. Burnham. 1999. Program MARK: survival estimation from populations of marked animals. Bird Study 46 Supplement:120-138.

Otis, D. L., and G. C. White. 1999. Autocorrelation of location estimates in the analysis of radiotracking data. Journal of Wildlife Management 63:1039-1044.

Conner, M. M., and G. C. White. 1999. Effects of individual heterogeneity in estimating the persistence of small populations. Natural Resources Modeling 12:109-127.

White, G. C., W. M. Block, J. P. Ward Jr., J. L. Ganey, A. B. Franklin, S. L. Spangle, J. R. Vahle, S. R. Rinkevich, W. H. Moir, F. P. Howe, and J. L. Dick Jr. 1999. Science Versus Political Reality in Delisting Criteria for a Threatened Species: The Mexican Spotted Owl Experience. Transactions of the 64th North American Wildlife and Natural Resources Conference 64:292-306.

Anderson, D. R., K. P. Burnham, A. B. Franklin, R. J. Gutiérrez, E. D. Foresman, R. G. Anthony, G. C. White, and T. M. Shenk. 1999. A protocol for conflict resolution in analyzing empirical data related to natural resource controversies. Wildlife Society Bulletin 27:1050-1058.

Boyce, M. S., A. R. E. Sinclair and G. C. White. 1999. Seasonal compensation of predation and harvesting. OIKOS 87:419-426.

White, G. C. 2000. Modeling Population Dynamics. Pages 84-107 in S. Demarais and P. R. Krausman, eds. Ecology and Management of Large Mammals in North America. Prentice-Hall, Upper Saddle River, New Jersey, USA.

White, G. C. 2000. Population viability analysis: data requirements and essential analyses. Pages 288-331 in L. Boitani and T. K. Fuller, editors. Research techniques in animal ecology: controversies and consequences. Columbia University Press, New York, New York, USA.

Bowden, D. C., G. C. White, and R. M. Bartmann. 2000. Optimal allocation of sampling effort for monitoring a harvested mule deer population. Journal of Wildlife Management 64:1013-1024.

White, G. C., D. J. Freddy, R. B. Gill, and J. H. Ellenberger. 2001. Effect of adult sex ratio on mule deer and elk productivity in Colorado. Journal of Wildlife Management 65:436-444.

Savarie, P. J., J. A. Shivik, G. C. White, J. C. Hurley, and L. Clark. 2001. Use of acetaminophen for large-scale control of Brown Treesnakes. Journal of Wildlife Management 65:356-365.

White, G. C. 2001. Why take calculus? Rigor in wildlife management. Wildlife Society Bulletin 29:380-386.

White, G. C. 2001. Statistical models: keys to understanding the natural world. Pages 35-56 in T. M. Shenk and A. B. Franklin, editors. Modeling in Natural Resource Management. Island Press, Washington, D. C.

Block, W. M., A. B. Franklin, J. P. Ward Jr., J. L. Ganey, and G. C. White. 2001. Design and implementation of monitoring studies to evaluate the success of ecological restoration on wildlife. Restoration Ecology 9:293-303.

Anderson, D. R., K. P. Burnham, and G. C. White. 2001. Kullback-Leibler information in resolving natural resource conflicts when definitive data exist. Wildlife Society Bulletin 29:1260-1270.

White, G. C., and T. M. Shenk. 2001. Population estimation with radio-marked animals. Pages 329-350 in J. J. Millspaugh and J. M. Marzluff, editors. Design and Analysis of Wildlife Radiotelemetry Studies. Academic Press, San Diego, California, USA.

Conner, M. M., G. C. White, and D. J. Freddy. 2001. Elk movement in response to early-season hunting in northwest Colorado. Journal of Wildlife Management 65:926-940.

White, G. C., K. P. Burnham, and D. R. Anderson. 2001. Advanced features of Program Mark. Pages 368-377 in R. Field, R. J. Warren, H. Okarma, and P. R. Sievert, editors. Wildlife, Land, and People: Priorities for the 21st Century. Proceedings of the Second International Wildlife Management Congress. The Wildlife Society, Bethesda, Maryland, USA.

Barker, R. J., and G. C. White. 2001. Joint analysis of live and dead encounters of marked animals. Pages 361-367 in R. Field, R. J. Warren, H. Okarma, and P. R. Sievert, editors. Wildlife, Land, and People: Priorities for the 21st Century. Proceedings of the Second International Wildlife Management Congress. The Wildlife Society, Bethesda, Maryland, USA.

White, G. C., A. B. Franklin, and T. M. Shenk. 2002. Estimating parameters of PVA models from data on marked animals. Pages 169-190 in S. R. Beissinger and D. R. McCullough, editors. Population Viability Analysis. University of Chicago Press, Chicago, Illinois, USA.

White, G. C., and B. C. Lubow. 2002. Fitting population models (Quattro) (Excel) to multiple sources of observed data. Journal of Wildlife Management 66:300-309.

Burnham, K. P. and G. C. White. 2002. Evaluation of some random effects methodology applicable to bird ringing data. Journal of Applied Statistics 29(1-4):245-264.

White, G. C. 2002. Discussant: The Use of Auxiliary Variables in Capture-Recapture Modeling: An Overview. Journal of Applied Statistics 29(1-4):103-106.

Otis, D. L. and G. C. White. 2002. Re-analysis of a banding study to test effects of an experimental increase in bag limits of mourning doves. Journal of Applied Statistics 29(1-4):479-495.

Dinsmore, S. J., G. C. White, and F. L. Knopf. 2002. Advanced techniques for modeling avian nest survival. Ecology 83:3476-3488.

Boulanger J., G. C. White, B. N. McLellan, J. Woods, M. F. Proctor, S. Himmer. 2003. A meta-analysis of grizzly bear DNA mark-recapture projects in British Columbia. Ursus 13:137-152.

Bowden, D. C., G. C. White, A. B. Franklin, and J. L. Ganey. 2003. Estimating population size with correlated sampling unit estimates. Journal of Wildlife Management 67:1-10.

Zablan, M. A., C. E. Braun, and G. C. White. 2003. Estimation of northern sage-grouse survival in North Park, Colorado. Journal of Wildlife Management 67:144-154.

Parmenter, R. R., T. L. Yates, D. R. Anderson, K. P. Burnham, J. L. Dunnum, A. B. Franklin, M. T. Friggens, B. C. Lubow, M. Miller, G. S. Olson, C. A. Parmenter, J. Pollard, E. Rexstad, T. M. Shenk, T. R. Stanley, and G. C. White. 2003. Small-mammal density estimation: a field comparison of grid-based vs. web-based density estimators. Ecological Monographs 73:1-26.

Dinsmore, S. J., G. C. White, and F. L. Knopf. 2003. Annual survival and population estimates of mountain plovers in southern Phillips County, Montana. Ecological Applications 13:1013-1026.

Vieira, M. E. P., M. M. Conner, G. C. White, and D. J. Freddy. 2003. Effects of archery hunter numbers and opening dates on elk movement. Journal of Wildlife Management. 67:717-728.

Phillips, G. E., and G. C. White. 2003. Pronghorn population response to coyote control: modeling and management. Wildlife Society Bulletin 31:1162-1175.

Reynolds, R. T., G. C. White, S. M. Joy, and R. W. Mannan. 2004. Effects of radiotransmitters on Northern Goshawks: do tail mounts lower survival of breeding males? Journal of Wildlife Management 68:25–32.

Franklin, A. B., R. J. Gutiérrez, J. D. Nichols, M. E. Seamans, G. C. White, G. S. Zimmerman, J. E. Hines, T. E. Munton, W. S. LaHaye, J. A. Blakesley, G. N. Steger, B. R. Noon, D. W. H. Shaw, J. J. Keane, T. L. McDonald, and S. Britting. 2004. Population dynamics of the California spotted owl (Strix occidentalis occidentalis): A meta-analysis. Ornithological Monograph 54:1–54.

Converse, S. J., B. G. Dickson, G. C. White, and W. M. Block. 2004. Estimating small mammal abundance on fuels treatment units in southwestern ponderosa pine forests. Pages 113–120 in van Riper III, C., and K. L. Cole, editors. The Colorado Plateau Cultural, Biological, and Physical Research. The University of Arizona Press, Tucson, Arizona, USA.

Peterson, D. P., K. D. Fausch, and G. C. White. 2004. Population ecology of an invasion: effects of brook trout on native cutthroat trout. Ecological Applications 14:754–772.

Norman, G. W., M. M. Conner, J. C. Pack, and G. C. White. 2004. Effects of fall hunting on survival of male wild turkeys in Virginia and West Virginia. Journal of Wildlife Management 68: 393–404.

Barker, R. J., K. P. Burnham, and G. C. White. 2004. Encounter history modeling of joint mark-recapture, tag-resighting and tag-recovery data under temporary emigration. Statistica Sinica 14:1037–1055.

Flanders-Wanner, B. L., G. C. White, and L. L. McDaniel. 2004. Weather and prairie grouse: dealing with effects beyond our control. Wildlife Society Bulletin 32: 22–34.

Walsh, D. P., G. C. White, T. E. Remington, and D. C. Bowden. 2004. Evaluation of the lek count index for greater sage-grouse. Wildlife Society Bulletin 32: 56–68.

Freddy, D. J., G. C. White, M. C. Kneeland, R. H. Kahn, J. W. Unsworth, W. J. deVergie, V. K. Grahm, J. H. Ellenberger, and C. H. Wagner. 2004. How many mule deer are there? Challenges of credibility in Colorado. Wildlife Society Bulletin 32:916–927.

Craig, G. R., G. C. White, and J. H. Enderson. 2004. Survival, recruitment, and rate of population change of the peregrine falcon population in Colorado. Journal of Wildlife Management 68:1032–1038.

Flanders-Wanner, B. L., G. C. White, and L. L. McDaniel. 2004. Validity of prairie grouse harvest-age ratios as production indices. Journal of Wildlife Management 68:1088–1094.

Ganey, J. L., G. C. White, D. C. Bowden, and A. B. Franklin. 2004. Evaluating methods for monitoring populations of Mexican spotted owls: a case study. Pages 337–385 in Thompson, W. L., editor. Sampling Rare or Elusive Species: Concepts, Designs and Techniques for Estimating Population Parameters. Island Press, Washington, D.C., USA.

Barker, R. J., and G. C. White. 2004. Towards the mother-of-all-models: customised construction of the mark recapture likelihood function. Animal Biodiversity and Conservation 27.1: 177–185.

Otis, D. L., and G. C. White. 2004. Evaluation of ultrastructure and random effects band recovery models for estimating relationships between survival and harvest rates in exploited populations. Animal Biodiversity and Conservation 27.1: 157–173.

White, G. C., and J. E. Hines. 2004. Computing and software. Animal Biodiversity and Conservation 27.1: 175–176.

Bishop, C. J., G. C. White, D. J. Freddy, and B. E. Watkins. 2005. Effect of limited antlered harvest on mule deer sex and age ratios. Wildlife Society Bulletin 33: 662–668.

White, G. C. 2005. Correcting wildlife counts with detection probabilities. Wildlife Research 32:211–216.

White, G. C., J. R. Dennis, and F. M. Pusateri. 2005. Area of black-tailed prairie dog colonies in eastern Colorado. Wildlife Society Bulletin 33:265–272.

Finley, D. J., G. C. White and J. P. Fitzgerald. 2005. Estimation of swift fox population size and occupancy rates in eastern Colorado. Journal of Wildlife Management 69:861–873.

Barker, R. J., G. C. White, and M. McDougal. 2005. Movement of paradise shelduck between molt sites: a joint multistate-dead recovery mark recapture model. Journal of Wildlife Management 69:1194–1201.

White, G. C., J. R. Dennis, and F. M. Pusateri. 2005. Response to: Overestimation bias in estimate of black-tailed prairie dog abundance in Colorado. Wildlife Society Bulletin 33:1452–1455.

Dinsmore, S. J., G. C. White, and F. L. Knopf. 2005. Mountain plover responses to black-tailed prairie dogs in Montana. Journal of Wildlife Management 69: 1546–1553.

Schwartz, C. C., M. A. Haroldson, G. C. White, R. B. Harris, S. Cherry, K. A. Keating, D. Moody, and C. Servheen. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the Greater Yellowstone Ecosystem. Wildlife Mongraphs 161.

Schwartz, C. C., M. A. Haroldson, and G. C. White. 2006. Study area and methods for collecting and analyzing demographic data on grizzly bears in the Greater Yellowstone Ecosystem. Pages 9–16 in Schwartz, C. C., M. A. Haroldson, G. C. White, R. B. Harris, S. Cherry, K. A. Keating, D. Moody, and C. Servheen. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the Greater Yellowstone Ecosystem. Wildlife Mongraphs 161.

Schwartz, C. C., M. A. Haroldson, and G. C. White. 2006. Survival of cub and yearling grizzly bears in the Greater Yellowstone Ecosystem. Pages 25–32 in Schwartz, C. C., M. A. Haroldson, G. C. White, R. B. Harris, S. Cherry, K. A. Keating, D. Moody, and C. Servheen. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the Greater Yellowstone Ecosystem. Wildlife Mongraphs 161.

Haroldson, M. A. C. C. Schwartz, and G. C. White. 2006. Survival of independent grizzly bears in the Greater Yellowstone Ecosystem. Pages 33–43 in Schwartz, C. C., M. A. Haroldson, G. C. White, R. B. Harris, S. Cherry, K. A. Keating, D. Moody, and C. Servheen. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the Greater Yellowstone Ecosystem. Wildlife Mongraphs 161.

Harris, R. B., C. C. Schwartz, M. A. Haroldson, and G. C. White. 2006. Trajectory of the Yellowstone grizzly bear population under alternative survival rates. Pages 44–56 in Schwartz, C. C., M. A. Haroldson, G. C. White, R. B. Harris, S. Cherry, K. A. Keating, D. Moody, and C. Servheen. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the Greater Yellowstone Ecosystem. Wildlife Mongraphs 161.

Anthony, R.G., E. D. Forsman, A. B. Franklin, D. R. Anderson, K. P. Burnham, G. C. White, C. J. Schwarz, J. D. Nichols, J. E. Hines, G. S. Olson, S. H. Ackers, L. S. Andrews, B. L. Biswell, P. C. Carlson, L. V Diller, K. M. Dugger, K. E. Fehring, T. L. Fleming, R. P. Gerhardt, S. A. Gremel, R. J. Gutiérrez, P. J. Happe, D. R. Herter, J. M. Higley, R. B. Horn, L. L. Irwin, P. J. Loschl, J. A. Reid, and S. G. Sovern. 2006. Status and trends in demography of northern spotted owls, 1985–2003. Wildlife Monographs 163:1–48.

Fields, T. L., G. C. White, W. C. Gilbert, and R. D. Rodgers. 2006. Lesser prairie-chicken nest and brood survival in west central Kansas. Journal of Wildlife Management 70:931-938.

McClintock, B. T., G. C. White, and K. P. Burnham. 2006. A robust design mark-resight abundance estimator allowing heterogeneity in resighting probabilities. Journal of Agricultural, Biological, and Ecological Statistics 11:231–248.

Saltz, D., D. I. Rubenstein, and G. C. White. 2006. The impact of increased environmental stochasticity due to climate change on the dynamics of Asiatic wild ass. Conservation Biology 20:1402-1409.

Converse, S. J., W. M. Block, and G. C. White. 2006. Small mammal population and habitat responses to forest thinning and prescribed fire. Forest Ecology and Management 228:263-273.

Franklin, A. B., J. D. Nichols, R. G. Anthony, K. P. Burnham, D. R. Anderson, G. C. White, and E. D. Forsman. 2006. Comment on “Are survival rates for northern spotted owls biased?” Canadian Journal of Zoology 84:1375–1379.

Converse, S. J., G. C. White, and W. M. Block. 2006. Small mammal responses to thinning and wildfire in ponderosa pine-dominated forests of the southwestern USA. Journal of Wildlife Management 70:1711–1722.

Converse, S. J., G. C. White, K. L. Farris, and S. Zack. 2006. Small mammal responses to forest fuel reduction: national scale results from the Fire and Fire Surrogate Project. Ecological Applications 16:1717–1729.

Mason, J. R., L. H. Carpenter, M. Cox, J. C. Devos, J. Fairchild, D.J. Freddy, J. R. Heffelfinger, R. H. Kahn, S. M. McCorquodale, D. F. Pac, D. Summers, G. C. White, and B. K. Williams. 2006. A case for standardized ungulate surveys and data management in the western United States. Wildlife Society Bulletin 34:1238–1242.

White, G. C., W. L. Kendall, and R. J. Barker. 2006. Multistate survival models and their extensions in program MARK. Journal of Wildlife Management 70:1521–1529.

McClintock, B. T., and G. C. White. 2007. Bighorn sheep abundance following a suspected pneumonia epidemic in Rocky Mountain National Park. Journal of Wildlife Management 71:183–189.

Pac, D. F., and G. C. White. 2007. Survival and cause-specific mortality of male mule deer under different hunting regulations in the Bridger Mountains, Montana. Journal of Wildlife Management 71:816–827.

Cherry, S., G. C. White, K. A. Keating, M. A. Haroldson, C. C. Schwartz. 2007. Evaluating estimators of the numbers of females with cubs-of-the-year in the Yellowstone grizzly bear population. Journal of Agricultural, Biological, and Ecological Statistics 12:195–215.

Heisey, D. M., T. L. Shaffer, and G. C. White. 2007. The ABCs of nest survival: theory and application from a biostatistical perspective. Studies in Avian Biology 34:13–33.

Bishop, C. J., D. J. Freddy, G. C. White, B. E. Watkins, T. R. Stephenson, and L. L. Wolfe. 2007. Using vaginal implant transmitters to aid in capture of neonates from marked mule deer. Journal of Wildlife Management 71:945–954.

Magle, S. R., B. T. McClintock, D. W. Tripp, G. C. White, M. F. Antolin, and K. R. Crooks. 2007. Mark-resight methodology for estimating population densities for prairie dogs. Journal of Wildlife Management 71:2067–2073.

Gragg, J. E., G. H. Rodda, J. A. Savidge, G. C. White, K. Dean-Bradley, and A. R. Ellingson. 2007. Response of brown treesnakes to reduction of their rodent prey. Journal of Wildlife Management 71:2311–2317.

Harris, R. L., G. C. White, C. C. Schwartz, and M. A. Haroldson. 2007. Population growth of Yellowstone grizzly bears: uncertainty and future monitoring. Ursus 18:168–178.

Bestgen, K. R., J. A. Hawkins, G. C. White, K. D. Chrisopherson, M. Hudson, M. H. Fuller, D. C. Kitcheyan, R. Brunson, P. Badame, G. B. Haines, J. Jackson, C. D. Walford, and T. A. Sorensen. 2007. Population status of Colorado pikeminnow in the Green River Basin, Utah and Colorado. Transactions of the American Fisheries Society 136:1356–1380.

Rogers, K. B., and G. C. White. 2007. Analysis of movement and habitat use from telemetry data. Pages ???–??? in Analysis and Interpretation of Freshwater Fisheries Data, American Fisheries Society, Bethesda, Maryland.

Martin, D. J., G. C. White, and F. M. Pusateri. 2007. Monitoring swift fox populations in eastern Colorado. Southwestern Naturalist 52:541–551.

White, G. C. 2008. Closed population estimation models and their extensions in program MARK. Environmental and Ecological Statistics 15:89-99.

Boulanger, J., G. C. White, M. Proctor, G. Stenhouse, G. MacHutcheon, and S. Himmer. 2008. Use of occupancy models to estimate the influence of previous live captures on DNA-based detection probabilities of grizzly bears. Journal of Wildlife Management 72:589-595.

Bishop, C. J., G. C. White, and P. M. Lukacs. 2008. Evaluating dependence among mule deer siblings in fetal and neonatal survival analyses. Journal of Wildlife Management 72:1085–1093.

Odell, E. A., F. M. Pusateri, and G. C. White. 2008. Estimation of occupied and unoccupied black-tailed prairie dog colony acreage in Colorado. Journal of Wildlife Management 72:1311–1317.

Huwer, S. L., D. R. Anderson, T. E. Remington, and G. C. White. 2008. Using human-imprinted chicks to evaluate the importance of forbs to sage-grouse. Journal of Wildlife Management 72:1622–1627.

Brand, L. A., G. C. White, and B. R. Noon. 2008. Factors influencing avian species richness and community composition of breeding birds in a desert riparian corridor. Condor 110:199–210.

McMahon, C. R., I. C. Field, C. J. A. Bradshaw, G. C. White, and M. A. Hindell. 2008. Tracking and data-logging devices attached to elephant seals do not affect individual mass gain or survival. Journal of Experimental Marine Biology and Ecology 360:71–77.

Schwartz, C. C., and G. C. White. 2008. Estimating reproductive rates for female bears: proportions versus transition probabilities. Ursus 19:1–12.

C. Moritz, C., J. L. Patton, C. J. Conroy, J. L. Parra, G. C. White, and S. R. Beissinger. 2008. Impact of a century of climate change on small mammal communities in Yosemite National Park. Science 322:261–264.

Conn, P. B., D. R. Diefenbach, J. L. Laake, M. A. Ternent, and G. C. White. 2008. Bayesian analysis of wildlife age-at-harvest data. Biometrics 64:1170–1177.

McClintock, B.T., White, G.C., Burnham, K.P. and Pryde, M.A. 2009. A generalized mixed effects model of abundance for mark-resight data when sampling is without replacement. Pages 271–289 in Modeling Demographic Processes in Marked Populations, D.L. Thomson, E.G. Cooch and M.J. Conroy, eds. Springer, New York, New York, USA.

Conn, P. B., G. C. White, and J. L. Laake. 2009. Simulation performance of Bayesian estimators of abundance employing age-at-harvest and mark-recovery data. Pages 965–986 in Modeling Demographic Processes in Marked Populations, D.L. Thomson, E.G. Cooch and M.J. Conroy, eds. Springer, New York, New York, USA.

White, G. C., K. P. Burnham, and R. J. Barker. 2009. Evaluation of a Bayesian MCMC random effects inference methodology for capture-mark-recapture data. Pages 1119–1127 in Modeling Demographic Processes in Marked Populations, D.L. Thomson, E.G. Cooch and M.J. Conroy, eds. Springer, New York, New York, USA.

Thomson, D. L., M. J. Conroy, D. R. Anderson, K. P. Burnham, E. G. Cooch, C. M. Francis, J.-D. Lebreton, M. S. Lindberg, B. J. T. Morgan, D. L. Otis, and G. C. White. 2009. Standardising terminology and notation for the analysis of demographic processes in marked populations. Pages 1099–1106 in Modeling Demographic Processes in Marked Populations, D.L. Thomson, E.G. Cooch and M.J. Conroy, eds. Springer, New York, New York, USA.

Kendall, K. C., J. B. Stetz, J. B. Boulanger, A. C. McLeod, D. Paetkau, and G. C. White. 2009. Demography and genetic structure of a recovering grizzly bear population. Journal of Wildlife Management 73:3–17.

Andelt, W. F., G. C. White, P. M. Schnurr, and K. W. Navo. 2009. Occupancy of random plots by white-tailed and Gunnison's prairie dogs. Journal of Wildlife Management 73:35–44.

Zimmerman, G. S., W. L. Kendall, T. J. Moser, G. C. White, and P. F. Doherty, Jr. 2009. Temporal patterns of apparent leg band retention in North American geese. Journal of Wildlife Management 73:82–88.

McClintock, B. T., G. C. White, M. F. Antolin, and D. W. Tripp. 2009. Estimating abundance using mark-resight when sampling is with replacement or the number of marked individuals is unknown. Biometrics 65:237–246.

Bears, H., K. Martin, and G. C. White. 2009. Breeding in high-elevation habitat results in shift to slower life history strategy within a single species. Journal of Animal Ecology 78:365–375.

McClintock, B. T., and G. C. White. 2009. A less field-intensive robust design for estimating demographic parameters with mark-resight data. Ecology 90:313–320.

McMahon, C. R., and G. C. White. 2009. Tag loss probabilities are not independent: assessing and quantifying the assumption of independent tag transition probabilities from direct observations. Journal of Experimental Marine Biology and Ecology 372:36–42.

Bishop, C. J., B. E. Watkins, L. L. Wolfe, D. J. Freddy, and G. C. White. 2009. Evaluating mule deer body condition using serum thyroid hormone concentrations. Journal of Wildlife Management 73:462–467.

Bishop, C. J., G. C. White, D. J. Freddy, B. E. Watkins, and T. R. Stephens. 2009. Effect of enhanced nutrition on mule deer population rate of change. Wildlife Monographs 172:1–28.

Zimmerman, G. S., T. J. Moser, W. L. Kendall, P. F. Doherty Jr., G. C. White, and D. F. Caswell. 2009. Factors influencing reporting and harvest probabilities in North American geese. Journal of Wildlife Management 73:710–719.

Lukacs, P. M., G. C. White, B. E.Watkins, R. H. Kahn, B. A. Banulis, D. J. Finley, A. A. Holland, J. A. Martens, and J. Vayhinger. 2009. Separating components of variation in survival of mule deer in Colorado. Journal of Wildlife Management 73:817–826.

Kendall, W. L., and G. C. White. 2009. A cautionary note on substituting spatial subunits for repeated temporal sampling in studies of site occupancy. Journal of Applied Ecology 46:1182–1188.

Sedinger, J. S., G. C. White, S. Espinosa, E. Partee, and C. E. Braun. 2010. Assessing compensatory versus additive harvest mortality: an example using Greater Sage-grouse Centrocercus urophasianus. Journal of Wildlife Management 74: 326–332.

McDonald, T. L., and G. C. White. 2010. A comparison of regression models for small counts. Journal of Wildlife Management 74:514–521.

Walsh, D. P., J. R. Stiver, G. C. White, T. E. Remington, and A. D. Apa. 2010. Population estimation techniques for lekking species. Journal of Wildlife Management 74:1607–1613.

Schwartz, C. C., M. A. Haroldson, and G. C. White. 2010. Hazards affecting grizzly bear survival in the Greater Yellowstone Ecosystem. Journal of Wildlife Management 74:654–667.

Zelasko, K. A., K. R. Bestgen, and G. C. White. 2010. Survival rates and movement of hatchery-reared razorback suckers in the upper Colorado River Basin, Utah and Colorado. Transactions of the American Fisheries Society 139:1478–1499.

Blakesley, J. A., M. E. Seamans, M. M. Conner, A. B. Franklin, G. C. White, R. J. Gutiérrez, J. E. Hines, J. D. Nichols, T. E. Munton, D. W. H. Shaw, J. J. Keane, G. N. Steger, and T. L. McDonald. 2010. Population dynamics of spotted owls in the Sierra Nevada, California. Wildlife Monographs 174:1–36.

Devineau, O., T. M. Shenk, G. C. White, P. F. Doherty Jr., P. M. Lukacs, and R. H. Kahn. 2010. Evaluating the Canada lynx reintroduction programme in Colorado: patterns in mortality. Journal of Applied Ecology 47:524–531.

Farnsworth, M. L., W. L. Kendall, P. F. Doherty Jr., R. S. Miller, G. C. White, J. D. Nichols, K. P. Burnham, and A. B. Franklin. 2011. Targeted surveillance for highly pathogenic avian influenza in migratory waterfowl across the United States. Pages 143–155 in S. K. Majundar, F. J. Brenner, J. E. Huffman, R. G. McLean, A. I. Panah, P. J. Pietrobon, S. P. Keeler, and S. E. Shive eds. Pandemic Influenza Viruses: Science, Surveillance and Public Health. The Pennsylvania Academy of Science, Easton, PA.

Forsman, E. D., R. G. Anthony, K. M. Dugger, E. M. Glenn, A. B. Franklin, G. C. White, C. J. Schwarz, K. P. Burnham, D. R. Anderson, J. D. Nichols, J. E. Hines, J. B. Lint, R. J. Davis, S. H. Ackers, L. S. Andrews, B. L. Biswell, P. C. Carlson, L. V. Diller, S. A.Gremel, D. R. Herter, J. M. Higley, R. B. Horn, J. A. Reid, J. Rockweit, J. Schaberel, T. J. Snetsinger, and S. G. Sovern. 2011. Population demography of northern spotted owls: 1985–2008. Studies in Avian Biology 40:1–103.

Saunders, W. C., K. D. Fausch, and G. C. White. 2011. Accurate estimation of salmonid abundance in small streams using three-pass removal electrofishing: an evaluation using marked fish. North American Journal of Fisheries Management 31:403–415.

Liebezeit, J.R., G.C. White, and, S. Zack. 2011. Breeding ecology of birds at Teshekpuk Lake: a key habitat site on the Coastal Plain of Alaska. Arctic 64:32–44.

Devineau, O., T. M. Shenk, P. F. Doherty Jr., G. C. White, and R. H. Kahn. 2011. Assessing release protocols used for Canada lynx reintroduction in Colorado. Journal of Wildlife Management 75:623–630.

Inman, R. M., M. L. Packila, K. H. Inman, A. J. McCue, G. C. White. J. Persson, B. C. Aber, M. Orme, K. L. Alt, S. L. Cain, J. A. Fredrick, B. Oakleak, and S. S. Sartorius. 2012. Spatial Ecology of wolverines at the southern periphery of distribution. Journal of Wildlife Management 76:778–792.

McClintock, B. T. and G. C. White. 2012. From NOREMARK to MARK: software for estimating demographic parameters using mark–resight methodology. Journal of Ornithology 152: S641–S650.

Doherty, Jr., P.F., G.C. White and K.P. Burnham. 2012. Comparison of model building strategies when all possible model structures cannot be constructed. Journal of Ornithology 152: S317–S323.

Pavlacky, D. C. Jr., J. A. Blakesley, G. C. White, D. J. Hanni, and P. M. Lukacs. 2012. Hierarchical multi-scale occupancy estimation for monitoring wildlife populations. Journal of Wildlife Management 76:154–162.

Kendall, W. L., G. C. White, J. E. Hines, C. A. Langtimm, and J. Yoshizaki . 2012. Estimating parameters of hidden Markov models based on marked individuals: use of robust design data. Ecology 93:913–920.

Jones, S. L., and G. C. White. 2012. The effect of habitat edges on nest survival of Sprague’s Pipits. The Wilson Journal of Ornithology 124:310–315.

Herendeen, D. L., and G. C. White. 2013. Statistical estimates of rare stamp populations. Pages 91–100 in T. Lera, J. H. Barwis and D. L. Herendeen, Editors. Proceedings of the First International Symposium on Analytical Methods in Philately. Smithsonian Contributions to History and Technology No. 57, Washington, D. C., USA.

Devineau, O., P. F. Doherty Jr., W. L. Kendall, T. M. Shenk, G. C. White, P. M. Lukacs, K. P. Burnham. 2013. Increased flexibility for modeling known-fate and nest-survival data using the multistate framework. Journal of Wildlife Management. In Press.

Ivan, J. S., G. C. White, and T. M. Shenk. 2013. Using auxiliary telemetry information to estimate animal density from capture-recapture data. Ecology 94:809–816.

Ivan, J. S., G. C. White, and T. M. Shenk. 2013. Using simulation to compare methods for estimating density from capture-recapture data. Ecology 94:809–816

Kendall, W. L., R. J. Barker, G. C. White, M. S. Lindberg, C. A. Langtimm and C. L. Peñaloza. 2013. Combining dead recovery, auxiliary observations and robust design data to estimate demographic parameters from marked individuals. Methods in Ecology and Evolution 4:828–835.

White, G. C., L. S. Cordes, and T. W. Arnold. 2013. Band reporting rates of waterfowl: does individual heterogeneity bias estimated survival rates? Ecology and Evolution.

Colorado Division of Wildlife Report to the Colorado Legislature: Declining Mule Deer Populations in Colorado: Reasons and Responses.

For questions or comments:

If you don't use WordPerfect, you may not have the WPMathA fonts used in some of the PDF documents provided above (which may result in various equations being unreadable). You can install the needed fonts from Corel by going to http://www.corel.com/6763/downloads/WordPerfect/wpwin/9/wpfonts.exe, which should solve the problem.

Faculty members in the Department of Fish, Wildlife & Conservation Biology of the College of Natural Resources at Colorado State University.

Last Modified 11/23/2013
Gary White / Gary.White at ColoState.edu