Sunil Kumar
Research Scientist, Natural Resource Ecology Laboratory
Affiliate Faculty, Department of Ecosystem Science and Sustainability
1499 Campus Delivery, B242 NESB, Colorado State University
Fort Collins, CO  80523-1499
Phone:(970) 491-7056, Fax:(970) 491-1965
e-mail: sunil.kumar 'at'

Current Research Interests:

Landscape ecology, niche modeling/species distribution modeling, biological invasions, pest risk analysis, geospatial analyses, remote sensing, GIS, spatial statistics, community ecology, climate change, and public health.

Peer-Reviewed Articles/Book Chapters/Abstracts (*indicates graduate student, **undergraduate student):

47. Kumar, S., Yee, W.L., and Neven, L.G. (In Press). Mapping global potential risk of establishment of Rhagoletis pomonella (Diptera: Tephritidae) using MaxEnt and CLIMEX niche models. Journal of Economic Entomology, DOI: 10.1093/jee/tow166.
46. Lestina**, J.T., Cook**, M., Kumar, S., Morisette, J.T., Ode, P., and Peairs, F. (In Press). MODIS imagery improves pest risk assessment: A case study of wheat stem sawfly (Cephus cinctus) in Colorado, USA. Environmental Entomology.
45. Galdino*, T.V.S., Kumar, S., Oliveira, L.S.S., Alfenas, A.C., Neven, L.G., Al-Sadi, A.M., and Picanco, M.C. (2016). Mapping global potential risk of mango sudden decline disease caused by fungus Ceratocystis fimbriata. PLoS ONE 11(7): e0159450. (PDF)
44. Sidder*, A.M., Kumar, S., Laituri, M., Sibold, J.S. (2016). Using spatiotemporal correlative niche models for evaluating the effects of climate change on mountain pine beetle. Ecosphere, 7(7):e01396. 10.1002/ecs2.1396. (PDF)
43. Javed, S.M.M., Raj, M., and Kumar, S. (In Press). Predicting potential habitat suitability for an endemic gecko Calodactylodes aureus and its conservation implications in India. Tropical Ecology.
42. Montecino, V., Molina, X., Bothwell, M., Muñoz, P., Carrevedo, M.L., Salinas, F., Kumar, S., Castillo*, M.L., Bizama*, G., and Bustamante, R.O. (2016). Spatio temporal population dynamics of the invasive diatom Didymosphenia geminata in central-southern Chilean rivers. Science of the Total Environment, 568: 1135-1145. (PDF)
41. West*, A.M., Kumar, S., Jarnevich, C.S. (2015). Regional Modeling of Large Wildfires Under Current and Potential Future Climates in Colorado and Wyoming, USA. Climate Change, 134: 565-577. (PDF)
40. Kumar, S., LeBrun, E.G., Stohlgren, T.J., Stabach*, J.A., McDonald, D.L., Oi, D.H., and LaPolla, J.S. (2015). Evidence of Niche Shift and Global Invasion Potential of Tawny Crazy Ant Nylanderia fulva. Ecology and Evolution, 5(20): 4628-4641. (PDF)
39. Jarnevich, C.S., Stohlgren, T.J., Kumar, S., Morisette, J.T., and Holcombe, T.R.(2015). Caveats for correlative species distribution modeling. Ecological Informatics 29: 6-15. (PDF)
38. Kumar, S., Neven, L.G., Zhu*, H., and Zhang, R.(2015). Assessing the Global Risk of Establishment of Cydia pomonella (Lepidoptera: Tortricidae) using CLIMEX and MaxEnt Niche Models. Journal of Economic Entomology, 108(4): 1708-1719. (PDF)
37. Strickland*, C., Dangelmayr, G., Shipman, P.D., Kumar, S., and Stohlgren, T.J. (2015). Network spread of invasive species and infectious diseases. Ecological Modelling 309-310: 1-9. (PDF)
36. West*, A.M., Kumar, S., Wakie, T., Brown, C.S., Stohlgren, T.J., Laituri, M., and Bromberg, J. (2015). Using high-resolution future climate scenarios to forecast Bromus tectorum invasion in Rocky Mountain National Park. PLoS ONE 10(2): e0117893. (PDF)
35. Kumar, S., Neven, L.G., and Wee, Y.L. (2014). Evaluating correlative and mechanistic niche models for assessing the risk of pest establishment. Ecosphere 5(7): 86. (PDF)
34. Montecino, V., Molina, X., Kumar, S., Castillo*, M.L.C., and Bustamante, R.O. (2014). Niche dynamics and potential geographic distribution of Didymosphenia geminata (Lyngbye) M. Schmidt, an invasive freshwater diatom in southern Chile. Aquatic Invasions 9(4): 507-519. (PDF)
33. Kumar, S., Neven, L.G., and Wee, Y.L. (2014). Assessing the Potential for Establishment of Western Cherry Fruit Fly Using Ecological Niche Modeling. Journal of Economic Entomology 107(3): 1032-1044. (PDF)
32. Kumar, S., Graham, J., West*, A.M., and Evangelista, P.H. (2014). Using district-level occurrences in MaxEnt for predicting the invasion potential of an exotic insect pest in India. Computers and Electronics in Agriculture, 103: 55-62. (PDF)
31. Kumar, S., Evangelista, P.H., Brown, C.S., Moore, J.C., Stohlgren, T.J., Graham, J.J., and Hanzlik, J.R. (2014). Recruiting and Retaining National Needs Fellows in the Area of Sciences for Agricultural Biosecurity. North American College Teacher Association (NACTA) Journal 58:66 (Peer-reviewed Abstract). (PDF); (Poster)
30. Schurich*, J.A., Kumar, S., Eisen, L., and Moore, C.G. (2014). Modeling Culex tarsalis abundance on the northern Colorado Front Range using a landscape-level approach. Journal of American Mosquito Control Association, 30: 7-20. (PDF)
29. Pfeifer, M., Lefebvre, V., Gardner, T., Arroyo-Rodriguez, V., Baeten, L., Banks-Leite, C., Barlow, J., Betts, M., Brunet, J., Cerezo, A., Cisneros, L., Collard, S., D’Cruze, N., da Silva Motta, C., Duguay, S., Eggermont, H., Eigenbrod, F., Hadley, A., Hanson, T., Hawes, J., Heartsill Scalley, T., Klingbeil, B., Kolb, A., Kormann, U., Kumar, S., Lachat, T., Lakeman Fraser, P., Lantschner, V., Laurance, W., Leal, I., Lens, L., Marsh, C., Medina-Rangel, G., Melles, S., Mezger, D., Oldekop, J., Overal, W., Owen, C., Peres, C., Phalan, B., Pidgeon, A., Pilia, O., Possingham, H., Possingham, M., Raheem, D., Ribeiro, D., Ribeiro Neto, J., Robinson, R., Robinson, W., Rytwinski, T., Scherber, C., Slade, E., Somarriba, E., Stouffer, P., Struebig, M., Tylianakis, J., Tscharntke, T., Tyre, A., Urbina Cardona, J., Vasconcelos, H., Wearn, O., Wells, K., Willig, M., Wood, E., Young, R., Bradley, A., and Ewers, R. (2014). BIOFRAG – A new database for analysing BIOdiversity responses to forest FRAGmentation. Ecology and Evolution, 4(9):1524-1537. (PDF)
28. Khanum*, K., Mumtaz, A.S., and Kumar, S., (2013). Predicting impacts of climate change on medicinal Asclepiads of Pakistan using Maxent Modeling. Acta Oecologica- International Journal of Ecology, 49: 23-31.(PDF)
27. Stohlgren, T.J., and Kumar, S. (2013). Endangered plants. In: Encyclopedia of Biodiversity, 2nd Edition. S. A. Levin (editor), Academic Press, Waltham, MA, Volume 3, pp. 205-215.(PDF)
26. Morain, S.A., Kumar, S., and Stohlgren, T.J. (2013). Environmental modeling for health. In: Environmental Tracking for Public Health Surveillance, Morain, S.A., and Budge, A.M. (editors), Taylor & Francis Group, Chapter 7, 293-332.(PDF)
25. Flory*, A. R., Kumar, S., Stohlgren, T. J., and Cryan, P. M. (2012). Environmental conditions associated with bat White-Nose Syndrome mortality in the north-eastern United States Journal of Applied Ecology, 49: 680-689.(PDF)
24. Nick*, Y., Stohlgren, T.J., Evangelista, P.H., Kumar, S., Graham, J., and Newman, G. (2012). Regional data refine local predictions: Modeling the distribution of plant species abundance on a portion of the Central Plains. Environmental Monitoring and Assessment, 184:5439-5451..(PDF)
23. Evangelista, P.H., Kumar, S., Stohlgren, T. J., and Nick, Y. (2011). Assessing forest vulnerability and the potential distribution of pine beetles under current and future climate scenarios in the Interior West of the U.S. Forest Ecology and Management, 262(3): 307-316.(PDF) (39 Citations; June 2016).
22. Evangelista, P., and Kumar. S. (2011) Trade and transportation is changing the game. Current Zoology 57(5): I-I (Guest Editorial).(PDF)
21. Bromberg*, J. E., Kumar, S., Brown, C. S., and Stohlgren, T. J. (2011). Distributional changes and range predictions of downy brome in Rocky Mountain National Park. Invasive Plant Science and Management, 4(2): 173-182.(PDF)
20. D'Cruze*, N. C., and Kumar, S. (2011). Effects of anthropogenic activities on lizard communities in northern Madagascar. Animal Conservation, 14(5): 542-552.(PDF)
19. Stohlgren, T. J., Kumar, S., Barnett, D., and Evangelista, P.H. (2011). Using maximum entropy modeling for optimal selection of sampling sites at regional scales. Diversity, 3: 252-261.(PDF)
18. York*, P., Evangelista, P.H., Kumar, S., Graham, J., Flather, C., and Stohlgren, T.J. (2011). A Habitat Overlap Analysis derived from Maxent for Tamarisk and the Southwestern Willow Flycatcher. Frontiers of Earth Science 5(2): 120-129.(PDF)
17. Stohlgren, T. J., Ma, P., Kumar, S., Rocca, M., Morisette, J. T., Jarnevich, C. S., and Benson, N. (2010). Ensemble habitat mapping of invasive plant species. Risk Analysis 30(2): 224-235.(PDF)
16. Lee*, A. T. K., Kumar, S., Brightsmith, D. J., and Marsden, S. J. (2010). Parrot claylick distribution in South America: do patterns of "where" help answer the question "why"? Ecography 32: 503-513.(PDF with Appendix)
15. Graham, J., Newman, G., Kumar, S., Jarnevich, C., Nick, Y., Crall, A., Stohlgren, T. J., and Evangelista, P. H. (2010). Bringing modeling to the masses: A web based system to predict potential species distributions. Future Internet, 2: 624-634.(PDF)
14. Evangelista, P. H., Stohlgren, T. J., Morisette, J. T., and Kumar, S. (2009). Mapping invasive tamarisk (Tamarix): a comparison of single-scene and time-series analyses of remotely sensed data. Remote Sensing 1: 519-533.(PDF)
13. Kumar, S., and Stohlgren, T. J.(2009). Maxent modeling for predicting suitable habitat for threatened and endangered tree Canacomyrica monticola in New Caledonia. Journal of Ecology and Natural Environment 1(4): 94-98.(PDF) (90 Citations; June 2016).
12. Kumar, S., Spaulding, S. A., Stohlgren, T. J., Hermann, K. A., Schmidt, T. S., and Bahls, L. L. (2009). Predicting habitat distribution for the freshwater diatom Didymosphenia geminata in the continental US. Frontiers in Ecology and the Environment 7(8): 415-420. (PDF with Appendix) (72 Citations; June 2016).
11. Kumar, S., Simonson, S. E. and Stohlgren, T. J. (2009). Effects of spatial heterogeneity on butterfly species richness in Rocky Mountain National Park, Colorado, USA. Biodiversity and Conservation 18(3): 739-763. (PDF) (23 Citations; June 2016).
10. D'Cruze*, N.C., Sabel, J.A., Dawson, J., Kumar, S. (2009). The influence of habitat type and structure on the abundance of Phelsuma madagascariensis grandis (Gekkonidae) in northern Madagascar. Herpetological Conservation and Biology 4(1): 55-61.(PDF)
9. D'Cruze*, N.C., Olsson, A., Henson, D., Kumar, S., and Emmett, D. (2009). The amphibians and reptiles of the Lower Onilahy River Valley, a temporary protected area in southwest Madagascar. Herpetological Conservation and Biology 4(1): 62-79.(PDF)
8. Evangelista, P.H., Kumar, S., Stohlgren, T. J., Jarnevich, C., Crall, A., Norman III, J. B., and Barnett, D. (2008). Modelling invasion for a habitat generalist and a specialist plant species. Diversity and Distributions 14: 808-817.(PDF) (84 Citations; June 2016).
7. Evangelista, P.H., Norman III, J. B., Berhanu, L, Kumar, S., and Alley, N. (2008). Predicting habitat suitability for endemic mountain nyala (Tragelaphus buxtoni) in Ethiopia. Wildlife Research 35: 409-416. (PDF)
6. Li, M.-Y., Ju, Y.-W., Kumar, S., and Stohlgren, T. J. (2008). Modeling potential habitat for Zebra mussel Dreissena polymorpha in the continental USA. Acta Ecologica Sinica (Chinese Ecology journal) 28(9): 4253-4258 .
5. Stohlgren, T. J., Jarnevich, C., Kumar, S. (2008). Forest legacies, climate change, altered disturbance regimes, invasive species, and water. Unasylva 229(58): 44-49.(PDF)
4. Evangelista, P.H., Kumar, S., Stohlgren, T. J., Newman, G., and Crall, A. (2007). Modeling aboveground biomass of Tamarix ramosissima in the Arkansas river basin in southeastern Colorado, USA. Western North American Naturalist 67(4): 503-509.(PDF)
3. Kumar, S., Stohlgren, T. J., and Chong, G. W. 2006. Spatial heterogeneity influences native and nonnative plant species richness. Ecology 87(12): 3186-3199.(PDF) (94 Citations; June 2016).
2. Lauenroth, W. K., Wade, A. A., Williamson, M.A., Ross, B.E., Kumar, S., and Cariveau, D. P. 2006. Uncertainty in calculations of net primary production for grasslands. Ecosystems 9: 843-851.(PDF)
1. Tiwari, A. K., Agarwal, A., Kumar, S., and Tiwari, S. C. 2005. Analysis of land use and biomass in Khanda watershed, Garhwal Himalaya, using satellite remote sensing data. Tropical Ecology 46(2): 253-263. (PDF)

*PDFs above are provided for non-commercial research and educational use; not for reproduction, distribution or commercial use.

Books, Thesis and Other Publications:

Book: Kumar, S. (2008). Effects of Spatial Heterogeneity on Species Richness: Methods to Quantify Spatial Heterogeneity and the Effects of Spatial Scale. 156 pp. VDM Verlag Dr. Muller, Saarbruken, Germany. ISBN: 978-3836488884. Available at
PhD Dissertation:"Effects of spatial heterogeneity on native and exotic plant and butterfly species richness in Rocky Mountain National Park, Colorado, USA".(Abstract). Major Advisor: Dr. Thomas J. Stohlgren (other committee members: Dr. Barry Noon, Dr. Michael Lefsky, and Dr. Hari Iyer).

Professional and Community Service:

  • Editorial Board, 'Tropical Ecology' (Oct. 2014 - present).
  • Editorial Board, 'Journal of Insect Science' (Jan. 2016 - present).
  • Editorial Board, 'Halteres' (Feb. 2015 - present)- A Peer Reviewed Open Access Research Journal in Entomology.
  • Guest editor, Current Zoology Special issue on Invasive Species (2010-2011).(PDF)
  • Editorial Board, 'Journal of Ecology and Natural Environment' (2009 - present).
  • Associate Editor, 'International Journal of Ecology and Development' (2012 - present).

  • Reviewer (# of proposals reviewed):
  • National Science Foundation (1): 2010
  • US-Israel Binational Science Foundation (1): 2011
  • Panelist USDA National Institute of Food and Agriculture (12): 2011
  • USDA Small Business Innovation Research (1): 2012
  • AgreenSkills Post-doc Fellowships (French National Institute for Agricultural Research) (2): 2014, 2015
  • Reviewer for >35 Journals (# of manuscripts reviewed): Biological Invasions (2), Landscape Ecology (3), Ecological Applications (4), Ecosphere (2), Journal of Ecology (1), Diversity and Distributions (1), Ecological Modelling (1), Ecology and Evolution (1), Global Ecology and Biogeography (1), PLOS ONE (3), Forest Ecology and Management (2), Computers and Electronics in Agriculture (1), Applied Vegetation Science (1), Aquatic Invasions (1), Entropy (1), Environmental Management (1), European Journal of Forest Research (1), Environmental Engineering Science (1), Rangeland Ecology and Management (1), Chinese Journal of Oceanology and Limnology (1), Nordic Journal of Botany (1), Invasive Plant Science and Management (1), Journal of Ecology and Natural Environment (3), Journal of Environmental Biology(1), Tropical Ecology (1), Conservation Biology (1), and Agroforestry Systems journals (1).
  • Teaching:

  • ESS330: Quantitative Reasoning in Ecosystem Science (Spring 2015)- Co-taught with Dr. Cameron Aldridge, Department of Ecosystem Science and Sustainability, Colorado State University (enrollement = 40).
  • ECOL592: Niche Models (2 credit seminar)- Co-taught with Dr. Barry Noon and Dr. Cameron Aldridge (Fall 2009, Spring 2011, Fall 2012, Fall 2013).
  • Watershed Problems course Spring 2014- Co-Taught, Dr. John Stednick, Department of Ecosystem Science and Sustainability, Colorado State University.
  • Teaching Assistant for ECOL592: Exploring Ecological Datasets (Fall 2007).
  • Guest Lecture (ECOL592) Notes:"Incorporating space in Ecological models- a lecture".(Download PDF).
  • Mathematics Tutor (1994-1996): taught mathematics to 11th and 12th grade students (2-3 batches of 8-12 students per academic year) .

  • Advising/Mentoring: Current

  • Ms. Jamie Fuller (Committee member), PhD.
  • Mr. Jung Gun Cho (Co-advisor), PhD.
  • Mr. Kevin McCartney (Advisor), MS, started Spring 2016.
  • Mr. Leo Franzone, Skills for Undergraduate Participation in Ecological Research (SUPER) program, Department of Ecosystem Science and Sustainability, CSU (started Nov. 2015).
  • Ms. Sara Hines, Skills for Undergraduate Participation in Ecological Research (SUPER) program, Department of Ecosystem Science and Sustainability, CSU (started Nov. 2015)
  • Advising/Mentoring: Past

  • Mr. Tarcisio V. S. Galdino (Advisor), PhD, Exchange Student from the University of Viçosa, Brazil (March, 2015- Jan. 2016).
  • Mr. Farajollah Tarnian (Advisor), PhD, Exchange Student from the University of Teheran, Iran (June-Dec.2015).
  • Mr. Jordan Lestina (Advisor), Undergraduate, Forestry/Forest Biology (Completed Dec. 2015), Honors Thesis: "Using MODIS Imagery for Pest Risk Assessment: A Case Study of Wheat Stem Sawfly (Cephus cinctus) in Colorado, USA".
  • Mr. Aaron Sidder (Co-advisor), MS, (Completed Summer 2015), Thesis: "Using spatiotemporal correlative niche models for evaluating the effects of climate change on mountain pine beetle" (PDF).
  • Ms. Amanda M. West (Co-advisor), PhD, (Completed Summer 2015), Dissertation: "Evaluating Risk for Current and Future 'Bromus tectorum' Invasion and Large Wildfires at Multiple Spatial Scales in Colorado and Wyoming, USA" (PDF); currently Post-Doc at Colorado State University
  • Mr. Anthony Vorster (Co-advisor), MS, (Completed Summer 2014), Thesis: "Relating severity of a mountain pine beetle outbreak to forest management history" (PDF); currently pursuing Ph.D. at Colorado State University
  • Ms. Jessica A. Schurich (Co-advisor), MS (Completed, Fall 2012), Thesis: "Modeling Culex tarsalis Coquillett abundance on the northern Colorado Front Range using a Landscape-level Approach" (PDF). Currently working with Colorado Mosquito Control Inc., Brighton, Colorado.

    h-index: 11.0 ( Researcher ID: A-6730-2009): Total citations- 641 (Senior author- 349)

    Google Scholar Profile: Sunil Kumar (h-index: 15.0; total citations: 1,145)

    ResearchGate: Sunil Kumar

    Curriculum Vitae

    Created on Septebmber 15, 2002
    Last modified on May, 2016, Sunil Kumar